An orphaned brontosaurus named Littlefoot sets off in search of the legendary Great Valley. A land of lush vegetation where the dinosaurs can thrive and live in peace. Along the way he meets four other young dinosaurs, each one a different species, and they encounter several obstacles as they learn to work together in order to survive.
A young mouse named Fievel and his family decide to migrate to America, a "land without cats," at the turn of the 20th century. But somehow, Fievel ends up in the New World alone and must fend off not only the felines he never thought he'd have to deal with again but also the loneliness of being away from home.
This time, while building a hideaway in their new home of the Great Valley, Littlefoot and the gang rescue a mysterious egg from two scheming egg-nappers and make a starling surprise - and new friend - when the egg hatches.
