The early years of the Madagascar heroes Alex the Lion, Marty the Zebra, Melman the Giraffe and Gloria the Hippo, as they grow up in a rescue habitat at the Central Park Zoo.
Eddie Garrick is a good-hearted man who has lost his belief in the wonder of Christmas. While spending time with his nine-year-old daughter Charlotte on Christmas Eve, he befriends a mysterious man in a red suit named Nick.
Apollo and Emma's love story is a fairy tale—until Emma mysteriously vanishes. Bereft, Apollo finds himself on a death-defying odyssey through a New York City he didn't know existed.
