Steve Coogan, an arrogant actor with low self-esteem and a complicated love life, is playing the eponymous role in an adaptation of "The Life and Opinions of Tristram Shandy, Gentleman" being filmed at a stately home. He constantly spars with actor Rob Brydon, who is playing Uncle Toby and believes his role to be of equal importance to Coogan's.
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To help build a family tree, single mom Amelia tracks down her daughter's biological father. It becomes a journey of trust, love, and discovering the meaning of family.
