Ten years after she was separated from her family, an eighteen-year-old orphan with vague memories of the past sets out to Paris in hopes of reuniting with her grandmother. She is accompanied by two con men, who intend to pass her off as the Grand Duchess Anastasia to the Dowager Empress for a reward.
A young man finds out that he holds the key to restoring hope and ensuring survival for the human race, while an alien species called the Drej are bent on mankind's destruction.
Born of a flower and growing to only a couple of inches tall, poor Thumbelina is worried she'll never meet someone her own size, until she happens to catch the eye of Prince Cornelius of the Fairies. Just as soon as she finds love, however, it's torn away from her when she is kidnapped by Ms. Toad. Now Thumbelina has to escape Ms. Toad's grasp and search for Prince Cornelius. Luckily, there's a whole city of animals willing to help her.
