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The story is set in the distant future. The land is ruined, and humanity establishes the mobile fort city Plantation. Pilots produced inside Plantation live in Mistilteinn, also know as the "birdcage." Children live there knowing nothing of the outside world or the freedom of the sky. Their lives consist of battling to carry out missions. Their enemies are mysterious giant lifeforms known as Kyouryuu, and the children pilot robots called Franxx to face off against them. For the children, riding the Franxx proves their existence. A boy named Hiro is called Code:016, and he was once known as a prodigy. However, he has fallen behind, and his existence seems unnecessary. Not piloting a Franxx is the same as ceasing to exist. One day, a mysterious girl known as "Zero Two" appears before him. Two horns grow out of her head.
Inspired by a small-statured pro volleyball player, Shouyou Hinata creates a volleyball team in his last year of middle school. Unfortunately the team is matched up against the "King of the Court" Tobio Kageyama's team in their first tournament and inevitably lose. After the crushing defeat, Hinata vows to surpass Kageyama. After entering high school, Hinata joins the volleyball team only to find that Tobio has also joined.
After a mysterious virus wipes out ninety percent of humanity, vampires emerge from the recesses of the earth to enslave mankind, treating them like livestock. After growing tired of donating blood in exchange for protection, 12-year-old Yuichiro Hyakuya and his best friend Mikaela Hyakuya plot an escape along with the other orphans. After being the only successful runaway, Yuichiro joins the Moon Demon Company, an extermination unit of the Japanese Imperial Demon Army dedicated to killing off vampires.
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