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It is the Taisho Period in Japan. Tanjiro, a kindhearted boy who sells charcoal for a living, finds his family slaughtered by a demon. To make matters worse, his younger sister Nezuko, the sole survivor, has been transformed into a demon herself. Though devastated by this grim reality, Tanjiro resolves to become a "demon slayer" so that he can turn his sister back into a human, and kill the demon that massacred his family.
A spy, an assassin and a telepath come together to pose as a family, each for their own reasons, while hiding their true identities from each other.
Denji has a simple dream—to live a happy and peaceful life, spending time with a girl he likes. This is a far cry from reality, however, as Denji is forced by the yakuza into killing devils in order to pay off his crushing debts. Using his pet devil Pochita as a weapon, he is ready to do anything for a bit of cash.
Dr. Jean Markham returns to the town she left as a teenager to take over her late father's medical practice. When a school-yard scuffle lands Charlie in her surgery, she invites him to visit the hives in her garden and tell his secrets to the bees, as she once did. The new friendship between the boy and the bee keeper brings his mother Lydia into Jean's world.
